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In 2015, a review recovered mDNA from historic canid specimens that were uncovered on Zhokhov Island along with the Yana river, arctic Siberia. These specimens included the mandible of the 360,000–400,000 YBP Canis cf. variabilis (the place cf. is a Latin phrase this means uncertain). Phylogenetic analyses of these canids unveiled 9 mDNA haplotyp

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